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Germania:

Most of these can also be applied to other Germanics, but
for the purpose of this post I’ll only be focusing on the similarities between
Germany and Germania.

Emotional stuntedness
+ awkwardness

Both Germania and Germany are unable to smile.
Well, at least expressively and regularly.

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Source: Hetarchive, Christmas Event 2010 + World Stars chapter 7. 

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Source: Hetarchive (Bamboo Thicket, Eng trans., Feb 24th, 2015). 

Perverted trope

While I’m not exactly favourable of this, there’s still the
running joke that Germans are closet perverts.

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Source: Volume 1 + Hetarchive (Bamboo Thicket, Eng trans., Feb 6th, 2015). 

Bridge between
nations: the organizer and ‘Mom of the group.’

Both bridge the gap between chaos and civility. In other
words, they organize those around them and keep them in line.

Ex: Germany demanding order at a world meeting vs Germania
making a snide comment when a meeting in Rome goes awry.

Source: Volume 1 + Hetarchive (Bamboo Thicket, Eng trans., May 3rd 2011).

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Ex: Both are adept planners.

Source: Hetarchive (Bamboo Thicket, Eng trans., Oct 20th 2013). 

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Ex: Both have an Italian to look after. This often includes
calming the latter down after an emotional outburst.

Source: Ibid + World Stars chapter 1

Rome + Germania

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Italy + Germany

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There’s also the tendency for the Italian to be late. 

Lastly, it’s obvious that they look pretty much identical
save for hair length.

Ancient Rome and the
Italy Brothers

Both brothers inherited the signature curl and a love for
women. Notice the positions of the curls and then compare them to Rome’s (i.e., one curl for each brother). 

Source: World Stars Chapter 20 + ibid 

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The split between them lies more with the expressions they
make.

Italy is the softer, more relaxed side of Rome, while Romano—while
also considerably vulnerable and soft—bears a more abrasive external shell; that is, when he’s not being a crybaby. 

The best way to go about this is to just show comparisons.
Notice how each Italy brother uniquely adopted specific expressions from Rome
that tailor well to their respective personalities. 

Source: Vol 1 + Vol 2

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Source: Vol 1

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Where did England’s tsundere personality come from?

It’s not a stereotype, so I would boil it down to creator preference.

What’s more, Himaruya has confirmed that Brittania (England’s mother) was a tsundere.

Source: Hetarchive (Bamboo Thicket, Eng trans., Sept 21rst, 2011). 

*Just want to clarify that I made a mistake in the past, back when I first began posting last year. Brittania doesn’t have an official appearance. The women surrounded by Rome in the We’re Shipwrecked strip haven’t been given official statuses as characters. Since Rome was known to always be surrounded by beautiful women, that’s what they remain as.*